Meikle, R.D. 1977: Flora of Cyprus 1
but accepted names differing
1Upper stem leaves not connate or amplexicaul, usually ternate; mericarps with prominent dorsal ridges
1'Upper steam leaves connate or amplexicaul, usually undivided and conspicuously different from the basal leaves; mericarps obscurely ridged
2Upper stem leaves alternate, amplexicaul with deeply cordate bases
2'Upper stem leaves opposite, connate along their basal margins