Bromus

Greuter, W. 1971: Bromus caroli-henrici, eine verkannte ostmediterrane Graminee. – Ann. Naturhist. Mus. Wien 75: 83-89Holmström, G. & Scholz, H. 2000: Bromus bidentatus spec. nova and B. lepidus, two odd brome-grasses (Gramineae). – Bot. Jahrb. Syst. 122: 195-200Meikle, R.D. 1985: Flora of Cyprus 2Scholz, H. 1974: Bromus chrysopogon Viviani: eine wieder als Art bewertete Graminee aus dem Mittelmeergebiet. – Willdenowia 7: 409-413Scholz, H. 1981: Bemerkungen über Bromus madritensis und B. rubens (Gramineae). – Willdenowia 11: 249-258Scholz, H. 1987: Delimiation and classification of Bromus fasciculatus (Poaceae). – Pl. Syst. Evol. 155: 277-282Scholz, H. 1995: Bromus regnii (Gramineae), a new endemic serpentine annual Brome-grass from Cyprus. – Willdenowia 25: 235-238Scholz, H. 2000: Bromus optimae, a new annual brome-grass (Gramineae) from Cyprus. – Bocconea 11: 85-87
This key is based on keys and data in the cited sources. It needs further testing, because several taxa are still controversial and their variability in and outside Cyprus is not fully understood.
1Spikelets oblong or wedge-shaped, not elliptic or lanceolate, nor tapering towards the tip; lower glume 1-nerved, the upper 3-nerved
1'Spikelets ovate, lanceolate, elliptic or oblong, tapering towards the tip; lower glume 3-7-nerved, upper 5-9-nerved
2Panicle loose, branches widely spreading, up to 10 cm long, carrying 1 or 2 spikelets, or sometime more
2'Panicle contracted or rather loose, but with the spikelets clustered in groups
3Spikelets 4-6 cm long (incl. awns), usually solitary at the tip of each branch
3'Spikelets 7-9 cm long (incl. awns), several on each branch
4Callus of the lemmas blunt, rounded
4'Callus of the lemmas sharply pointed
5Spikelets 7-9 cm long (incl. awns)
5'Spikelets less than 7 cm long (incl. awns), often much less
6Callus of the lemmas blunt, rounded
6'Callus of the lemmas sharply pointed
7Culms hairy below the inflorescence
7'Culms glabrous or very sparsely hairy
8Inflorescence broadly obovate, not acutely cuneate at the base; ligule up to 5 mm long, lacerate, milky-white; spikelets compressed; lemmas 13-15 x 2-2.5 mm, with a definite scarious margin; anthers 0.6-1 mm long; awns and lemmas straight
8'Inflorescence obovate, sharply cuneate at the base; ligule 3 mm long, hyaline; lemmas 9-16 x 1.5-2 mm, rounded on the back, with involute margins, which are hardly scarious; anthers 0.3-0.4 mm long; awns and lemmas curved outwards
9Spikelets 1.5-2.5 cm long; basal boundary line of lemma callus parabolic; abscission zone at rhachilla internode ovate; lower sheaths with dense villose hairs up to 0.7 mm long
9'Spikelets 2-3.5 cm long; basal boundary line of lemma callus semiorbicular; abscission zone at rhachilla internode orbicular; lower sheaths with dense to sparse short hairs
10Spikelets 3.5-6 cm long (incl. awns), sometimes even longer, erect, often dense, obovate; lemmas 12-19 x 3 mm; anthers 0.5-1 mm long
10'Spikelets up to 3 cm long (incl. awns), if slightly longer (B. tectorum) then the spikelets strongly compressed, with a tendency to turn to one side in clusters of parallel spikelets
11Lower sheaths with dense indumentum of short or long hairs; spikelets (excl. awns) 15-60 mm long
11'Lower sheaths with scattered patent hairs; spikelets (excl. awns) 15-25 mm long
12Panicle dense or somewhat loose, not flabellate or cuneate at the base; spikelets in parallel or roughly parallel clusters, erect or turned to one side; lemmas with silvery margins; anthers more than 0.3-0.4 mm long
12'Panicle very dense, flabellate, cuneate at base; lemmas 1.5-2 mm wide with involute margins, slightly recurved from the rhachilla; anthers 0.3-0.4 mm long
13Spikelets 1.5-2.5 cm long; basal boundary line of lemma callus parabolic; abscission zone at rhachilla internode ovate; lower sheaths with dense villose hairs up to 0.7 mm long
13'Spikelets 2-3.5 cm long; basal boundary line of lemma callus semiorbicular; abscission zone at rhachilla internode orbicular; lower sheaths with dense to sparse short hairs
14Glumes and lemmas rounded on the dorsal surface
14'Glumes and lemmas strongly keeled
15Awn of lemma inserted at the base of the notch
15'Awn of lemma inserted below the base of the notch
16Anthers (2.5-)3-5 mm long
16'Anthers 0.3-1.5(-2) mm long
17Panicle dense to rather loose, pedicels equalling or shorter than their spikelets; lemmas with prominent nerves
17'Panicle lax, most pedicels longer than their spikelets; lemmas with obscure nerves
18Lemmas with obtuse, sometimes indistinct apical teeth; awns 6-10 mm long, emerging up to 2 mm below apex of lemma
18'Lemmas with acute, distinct apical teeth; awns 12-14 mm long (6-8 on the lowest lemma), emerging 3-4 mm below apex of lemma
19Spikelets 3-5 cm long (incl. awns; often shorter in starved specimens of B. squarrosus)
19'Spikelets less than 3 cm long (incl. awns)
20Panicle oblanceolate or oblong in outline; spikelets terete, short-pedicellate, ascending; awns recurved at maturity
20'Panicle with spreading branches; spikelets laterally compressed
21Lemmas shortly bidentate
21'Lemmas biaristulate
22Lemmas 8-10 x 6 mm, with a broad hyaline margin, conspicuously widened above the middle, with the margins forming an obtuse angle; awns of the lower lemmas frequently reduced
22'Lemmas 11-14 x 5-6 mm, with a broad hyaline margin, not as above; lower lemmas with perfect awns, and 2 acuminate apical teeth; awns recurved at maturity
23Lemmas rhomboidal when flattened; margins conspicuously hyaline
23'Lemmas elliptic or oblanceolate when flattened; margins not conspicuously hyaline
24Spikelets hairy, hairs patent, long (c. 0.75 mm) and short, curved
24'Spikelets glabrous, or, if hairy, then the hairs usually equal, adpressed and short (usually not longer than 0.5 mm)
25Inflorescence a rather loose, nodding panicle; spikelets long-pedicellate, narrowly lanceolate; awns of the uppermost lemmas much longer than those of the lower
25'Inflorescence and spikelets not as above; the inflorescence erect, dense or rather loose
26Inflorescence very dense, obovate or oblong; lemmas 9-11 mm long; anthers 0.5 mm long
26'Inflorescence loose or somewhat dense; lemmas 8-9 mm long; anthers 1-2 mm long